I can start my 1999 crown Victoria with spark plug #1 disconnected but, if I connect it, it will shut off and will blow the fuse

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Question moved to model category.

Did you disconnect the battery recently before this happened? Is the anti theft light flashing?

https://www.google.com/search?q=1999+Ford+Crown+Victoria.+Cranks%2C+no+start

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You'll first need to find out if you are lacking fuel or spark to know where to go next. Do a basic test for fuel pressure with a gauge and spark at the plugs when it fails to start.

Owners Manual is on the internet

Google it

No fuse or relay will fix your issues,I doubt

It is doing its job(blowing when there is more current flowing than there should be). What circuit is that fuse in? Whatever the load is(light switch, radio, etc) has an internal short or the wiring between the fuse and the load is shorted to ground. Also never replace the fuse with one that has a higher current rating. That's when you start melting wires and other things that you don't want to melt. Good luck!

It could be loose or broken cables check terminal connections from alternator and also connections to battery

with out knowing what you have for tools unplug different parts of the system and try another fuse like the compressor clutch the pressure switches and plug them back in one at a time til the fuse blows if you have a ohm meter check to see if any part is shorted to ground

The reason a fuse blowing may be because of a short circuit. And you should check any wire connectors that plug into the pc board.

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Fix your car right away by clicking the link I attached
Ford Crown Victoria 1998 2012 Workshop Service repair manual Download

Car not starting or turning over indicates towards multiple possibilities. It can be issue with battery or weak battery. But if battery checks out ok, then it can be starter issue or faulty alternator or problem with low fuel pressure or no spark at spark plugs due to dirty spark plugs or faulty ignition coil or injectors getting cracked or dirty and needs to be cleaned.

To confirm the problem, the basic troubleshooting is required.

Click this link below and follow the troubleshooting:------

http://technoanswers.blogspot.in/2012/02/car-starting-issues.html

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This will help.
Thanks.

Attempt to change all of the spark plug wiring and spark plugs. Then start the vehicle.